* He studied medicine at the University of Göttingen, receiving his doctorate in 1866. His doctoral thesis was on the subject of *Botulism*. He didn't have a specific undergraduate degree before this; the German system at the time was different than the current model of separate undergraduate and graduate studies. His studies included practical training in hospitals and other medical settings.
While his formal education was primarily in medicine, his later achievements in bacteriology involved significant self-directed learning and research. His contributions were groundbreaking, even though they weren't gained through formal post-graduate training in the field of microbiology, which was still developing at the time.